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Spencer's biggest rival is her older sister, Melissa, and the two are constantly at odds.
Their relationship is often strained by the fact that Spencer has a habit of crushing on, and later kissing, Melissa's boyfriends (first Ian, then Wren).
Most of Aria's story line revolves on her love life. In season 5B, she deals with the fact that Alison is 'A', but later in 5B it's revealed that she isn't.
In season 6, she deals with her time in the dollhouse and Charlotte.
Ruling the group with an iron fist, the vicious but charismatic Ali went missing during a slumber party at the end of summer.
When Aria, Hanna, Spencer, and Emily begin receiving threatening messages from "A" a year after Ali goes missing, they think she is alive and is playing a trick on them.
Emily was closest to Alison before her disappearance and was also in love with her. After being rescued from A's dollhouse, she begins bonding with another victim of 'A': Sara Harvey , who was kept down there for 2 years.
She enjoys bullying Lucas, much to Hanna's annoyance.
After the appearance of A, Mona notices that Hanna is now acting strangely, and feels jealous that Hanna has rekindled her friendships with Aria, Emily, and Spencer.
From episode 2 of season 6, Alison finds a new love interest in Toby's police partney Lorenzo .
Mona Vanderwaal After Alison's disappearance, Mona completely transformed herself and befriended Hanna, and they both became popular students.